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Publications - John A. Byers
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1. Byers, J.A., Brewer, J.W., and Denna, D.W. 1976. Plant growth hormones in
pinyon insect galls. Marcellia 39:125-134.
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2. Byers, J.A., Wood, D.L., Browne, L.E., Fish, R.H., Piatek, B., and Hendry,
L.B. 1979. Relationship between a host plant compound, myrcene and
pheromone production in the bark beetle, Ips paraconfusus. Journal
of Insect Physiology 25:477-482.
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3. Byers, J.A. and Wood, D.L. 1980. Interspecific inhibition of the response
of the bark beetles, Dendroctonus brevicomis and Ips paraconfusus,
to their pheromones in the field. Journal of Chemical Ecology
6:149-164.
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4. Byers, J.A., Svihra, P., and Koehler, C.S. 1980. Attraction of elm bark
beetles to cut limbs on elm. Journal of Arboriculture
6:245-246. 
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5. Hendry, L.B., Piatek, B., Browne, L.E., Wood, D.L., Byers, J.A., Fish,
R.H., and Hicks, R.A. 1980. In vivo conversion of a labelled host
plant chemical to pheromones of the bark beetle Ips paraconfusus.
Nature 284:485.
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6. Byers, J.A. 1981a. Pheromone biosynthesis in the bark beetle, Ips
paraconfusus, during feeding or exposure to vapours of host plant
precursors. Insect Biochemistry 11:563-569.
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7. Byers, J.A. 1981b. Effect of mating on terminating aggregation during
host colonization in the bark beetle, Ips paraconfusus. Journal of
Chemical Ecology 7:1135-1147.
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8. Byers, J.A. 1981c. Versatile electronic timer for synchronous switching
of multiple electrical devices. Behavior Research Methods and
Instrumentation 13:381-383.
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9. Byers, J.A. and Wood, D.L. 1981a. Interspecific effects of pheromones on
the attraction of the bark beetles, Dendroctonus brevicomis and Ips
paraconfusus in the laboratory. Journal of Chemical Ecology
7:9-18.
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10. Byers, J.A. and Wood, D.L. 1981b. Antibiotic-induced inhibition of
pheromone synthesis in a bark beetle. Science 213:763-764.
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11. Byers, J.A. 1982. Male-specific conversion of the host plant compound,
myrcene, to the pheromone, (+)-ipsdienol, in the bark beetle,
Dendroctonus brevicomis. Journal of Chemical Ecology
8:363-371.
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12. Byers, J.A., and Poinar, G.O., Jr. 1982. Location of insect hosts by the
nematode, Neoaplectana carpocapsae, in response to temperature.
Behaviour 79:1-10.
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13. Byers, J.A. 1983a. Influence of sex, maturity and host substances on
pheromones in the guts of the bark beetles, Ips paraconfusus and
Dendroctonus brevicomis. Journal of Insect Physiology
29:5-13.
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14. Byers, J.A. 1983b. Bark beetle conversion of a plant compound to a
sex-specific inhibitor of pheromone attraction. Science
220:624-626.
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15. Byers, J.A. 1983c. Sex-specific responses to aggregation pheromone:
Regulation of colonization density in the bark beetle Ips
paraconfusus. Journal of Chemical Ecology 9:129-142.
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16. Byers, J.A. 1983d. Electronic fraction collector used for insect sampling
in the photoperiod-induced diel emergence of bark beetles.
Physiological Entomology 8:133-138.
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17. Byers, J.A., and Unkrich, M.A. 1983. Electronic light intensity control to
simulate dusk and dawn conditions. Annals of the Entomological
Society of America 76:556-558.
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18. Byers, J.A. 1984a. Electronic multiprobe thermometer and multiplexer for
recording temperatures of microenvironments in the forest litter
habitat of bark beetles (Coleoptera: Scolytidae). Environmental
Entomology 13:863-867.
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19. Byers, J.A. 1984b. Nearest neighbor analysis and simulation of
distribution patterns indicates an attack spacing mechanism in the
bark beetle, Ips typographus (Coleoptera: Scolytidae). Environmental
Entomology 13:1191-1200.
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20. Byers, J.A., Wood, D.L., Craig, J. and Hendry, L.B. 1984. Attractive and
inhibitory pheromones produced in the bark beetle, Dendroctonus
brevicomis, during host colonization: Regulation of inter- and
intraspecific competition. Journal of Chemical Ecology
10:861-877.
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21. Byers, J.A., Lanne, B.S., Löfqvist, J., Schlyter, F. and Bergström, G.
1985. Olfactory recognition of host-tree susceptibility by pine
shoot beetles. Naturwissenschaften 72:324-326.
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22. Byers, J.A. 1987a. Interactions of pheromone component odor plumes of
western pine beetle. Journal of Chemical Ecology
13:2143-2157.
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23. Byers, J.A. 1987b. Novel fraction collector for studying the oviposition
rhythm in the turnip moth. Chronobiology International
4:189-194.
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24. Lanne, B.S., Schlyter, F., Byers, J.A., Löfqvist, J., Leufvén, A.,
Bergström, G., Van Der Pers, J.N.C., Unelius, R., Baeckström, P. and
Norin, T. 1987. Differences in attraction to semiochemicals present
in sympatric pine shoot beetles, Tomicus minor and T. piniperda.
Journal of Chemical Ecology 13:1045-1067.
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25. Schlyter, F., Byers, J.A. and Löfqvist, J. 1987a. Attraction to pheromone
sources of different quantity, quality, and spacing:
Density-regulation mechanisms in bark beetle Ips typographus.
Journal of Chemical Ecology 13:1503-1523.
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26. Schlyter, F., Löfqvist, J. and Byers, J.A. 1987b. Behavioural sequence in
attraction of the bark beetle Ips typographus to pheromone sources.
Physiological Entomology 12:185-196.
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27. Schlyter, F., Birgersson, G., Byers, J.A., Löfqvist, J. and Bergström, G.
1987c. Field response of the spruce bark beetle, Ips typographus,
to aggregation pheromone candidates. Journal of Chemical Ecology
13:701-716.
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28. Byers, J.A. 1988a. Upwind flight orientation to pheromone in western pine
beetle tested with rotating windvane traps. Journal of Chemical
Ecology 14:189-198.
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29. Byers, J.A. 1988b. Novel diffusion-dilution method for release of
semiochemicals: Testing pheromone component ratios on western pine
beetle. Journal of Chemical Ecology 14:199-212.
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30. Byers, J.A., Birgersson, G., Löfqvist, J., and Bergström, G. 1988.
Synergistic pheromones and monoterpenes enable aggregation and host
recognition by a bark beetle, Pityogenes chalcographus.
Naturwissenschaften 75:153-155.
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31. Schlyter, F., Byers, J.A., Löfqvist, J., Leufvén, A., and Birgersson, G.
1988. Reduction of attack density of the bark beetles Ips
typographus and Tomicus piniperda on host bark by verbenone
inhibition of attraction to pheromone and host kairomone. pp. 53-68.
In: Payne, T.L. and Saarenmaa, H. (eds.), Integrated Control of
Scolytid Bark Beetles, Virginia Tech. Press, Blacksburg.
355 pp.
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32. Byers, J.A. 1989a. Chemical ecology of bark beetles. Experientia 45:271-
283.
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33. Byers, J.A. 1989b. Behavioral mechanisms involved in reducing competition
in bark beetles. Holarctic Ecology 12:466-476.
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34. Byers, J.A. and Löfqvist, J. 1989. Flight initiation and survival in the
bark beetle Ips typographus (Coleoptera: Scolytidae) during the
spring dispersal. Holarctic Ecology 12:432-440.
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35. Byers, J.A., Anderbrant, O., and Löfqvist, J. 1989a. Effective attraction
radius: A method for comparing species attractants and determining
densities of flying insects. Journal of Chemical Ecology
15:749-765.
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36. Byers, J.A., Högberg, H.E., Unelius, R., Birgersson, G. and Löfqvist, J.
1989b. Structure-activity studies on aggregation pheromone
components of Pityogenes chalcographus (Coleoptera: Scolytidae): all
stereoisomers of chalcogran and methyl 2,4-decadienoate. Journal of
Chemical Ecology 15:685-695.
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37. Byers, J.A., Lanne, B.S., and Löfqvist, J. 1989c. Host-tree unsuitability
recognized by pine shoot beetles in flight. Experientia
45:489-492.
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38. Birgersson, G., Byers, J.A., Bergström, G., and Löfqvist, J. 1990.
Production of pheromone components, chalcogran and methyl (E,Z)-2,4-
decadienoate, in the spruce engraver Pityogenes chalcographus.
Journal of Insect Physiology 36:391-395.
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39. Byers, J.A., and Birgersson, G. 1990. Pheromone production in a bark beetle
independent of myrcene precursor in host pine species.
Naturwissenschaften 77:385-387.
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40. Byers, J.A., Birgersson, G., Löfqvist, J., Appelgren, M., and Bergström,
G. 1990a. Isolation of pheromone synergists of bark beetle,
Pityogenes chalcographus, from complex insect-plant odors by
fractionation and subtractive-combination bioassay. Journal of
Chemical Ecology 16:861-876.
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41. Byers, J.A., Schlyter, F., Birgersson, G., and Francke, W. 1990b. E-
Myrcenol in Ips duplicatus: An aggregation pheromone component new
for bark beetles. Experientia 46:1209-1211.
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42. Byers, J.A. 1991a. Simulation of mate-finding behaviour of pine shoot
beetles, Tomicus piniperda. Animal Behaviour 41:649-660.
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43. Byers, J.A. 1991b. BASIC algorithms for random sampling and treatment
randomization. Computers in Biology and Medicine 21:69-77.
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44. Byers, J.A. 1991c. Cardcat: A scientific reference manager for IBM-
compatible personal computers. American Entomologist
37:240-242.
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45. Byers, J.A. 1991d. Pheromones and chemical ecology of locusts. Biological
Reviews 66:347-378.
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46. Byers, J.A. 1992a. Grid cell contour mapping of point densities: bark
beetle attacks, fallen pine shoots, and infested trees. Oikos
63:233-243.
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47. Byers, J.A. 1992b. Ref-list: Program to list references found in DOS text
files of scientific manuscripts. Computer Applications in the
Biosciences 8:196-198.
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48. Byers, J.A. 1992c. Dirichlet tessellation of bark beetle spatial attack
points. Journal of Animal Ecology 61:759-768.
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49. Byers, J.A. 1992d. Trends in chemical ecology revealed with a personal
computer program for searching data bases of scientific references
and abstracts. Journal of Chemical Ecology 18:1481-1495.
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50. Byers, J.A. 1992e. Optimal fractionation and bioassay plans for isolation
of synergistic chemicals: the subtractive-combination method.
Journal of Chemical Ecology 18:1603-1621.
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51. Byers, J.A. 1992f. Attraction of bark beetles, Tomicus piniperda,
Hylurgops palliatus, and Trypodendron domesticum and other insects
to short-chain alcohols and monoterpenes. Journal of Chemical
Ecology 18:2385-2402.
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52. Schlyter, F., Birgersson, G., Byers, J.A., and Bakke, A. 1992. The
aggregation pheromone of Ips duplicatus and its role in competitive
interactions with I. typographus (Coleoptera: Scolytidae).
Chemoecology 3:103-112.
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53. Zhang, Q.H., Byers, J.A., and Schlyter, F. 1992. Optimal attack density in
the larch bark beetle, Ips cembrae (Coleoptera: Scolytidae). Journal
of Applied Ecology 29:672-678.
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54. Byers, J.A. 1993a. Randomization algorithms in BASIC for experimental
design. Computers in Biology and Medicine 23:167-176.
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55. Byers, J.A. 1993b. Avoidance of competition by spruce bark beetles, Ips
typographus and Pityogenes chalcographus. Experientia 49:272-275.

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56. Byers, J.A. 1993c. Simulation and equation models of insect population
control by pheromone-baited traps. Journal of Chemical Ecology
19:1939-1956.
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57. Byers, J.A. 1993d. Orientation of bark beetles Pityogenes chalcographus
and Ips typographus to pheromone-baited puddle traps placed in
grids: A new trap for control of scolytids. Journal of Chemical
Ecology 19:2297-2316.
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58. Byers, J.A. 1993e. Conversion of BIOSIS Previews database files to
reference text files and reprint request forms. American
Entomologist 39:245-248.
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59. Zhang, Q.H., Byers, J.A., and Zhang, X.D. 1993. Influence of bark
thickness, trunk diameter and height on reproduction of the long-
horned beetle, Monochamus sutor (Col., Cerambycidae) in burned larch
and pine. Journal of Applied Entomology 115:145-154.
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60. Byers, J.A. 1995. Host tree chemistry affecting colonization in bark
beetles, in R.T. Cardé and W.J. Bell (eds.). Chemical Ecology of
Insects 2. Chapman and Hall, New York, pp. 154-213.
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61. Zolubas, P. and Byers, J.A., 1995. Recapture of dispersing bark beetles
Ips typographus L. (Col., Scolytidae) in pheromone-baited traps.
Journal of Applied Entomology 19:285-289.
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62. Byers, J.A. 1996a. Random selection algorithms for spatial and temporal
sampling. Computers in Biology and Medicine 26:41-52.
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63. Byers, J.A. 1996b. An encounter rate model of bark beetle populations
searching at random for susceptible host trees. Ecological Modelling
91:57-66.
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64. Byers, J.A. 1996c. Taxokey: a universal taxonomic key program using DOS
text files and graphics. Computer Applications in the Biosciences
12:185-189.
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65. Byers, J.A. 1996d. Correct calculation of Dirichlet polygon areas.
Journal of Animal Ecology 65:528-529.
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66. Byers, J.A. 1996e. Temporal clumping of bark beetle arrival at pheromone
traps: Modeling anemotaxis in chaotic plumes. Journal of Chemical
Ecology 22:2133-2155.
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67. Byers, J.A., Zhang, Q.H., Schlyter, F., and Birgersson, G. 1998.
Volatiles from nonhost birch trees inhibit pheromone response in
spruce bark beetles. Naturwissenschaften 85:557-561.
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68. Byers, J.A. 1999a. Database program to manage slides and images for
teaching and presentations. Educational Media International 36:77-80.
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69. Byers, J.A. 1999b. Flash cards and animation software for education.
Educational Media International 36:164-167.
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70. Byers, J.A. 1999c. Interactive learning using expert system quizzes on
the internet. Educational Media International 36:191-194.
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71. Byers, J.A. 1999d. Effects of attraction radius and flight paths on catch
of scolytid beetles dispersing outward through rings of pheromone
traps. Journal of Chemical Ecology 25:985-1005.
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72. Byers, J.A. 2000. Wind-aided dispersal of simulated bark beetles flying
through forests. Ecological Modelling 125:231-243.
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73. Byers, J.A., Zhang, Q.H., and Birgersson, G. 2000. Strategies of a bark
beetle, Pityogenes bidentatus, in an olfactory landscape.
Naturwissenschaften 87:503-507.
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74. El-Sayed, A.M. and Byers, J.A. 2000. Inhibitory effect of monoterpenes on
response of Pityogenes bidentatus to aggregation pheromone released
by piezoelectric sprayer for precision release of semiochemicals.
Journal of Chemical Ecology 26:1795-1809.
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75. Schlyter F., Zhang Q-H., Anderson P., Byers J.A., Wadhams L.J., Löfqvist,
J., and Birgersson G. 2000. Electrophysiological and behavioural responses
of Scandinavian Tomicus piniperda and T. minor (Coleoptera: Scolytidae),
to non-host leaf and bark volatiles. Canadian Entomologist 132:965-981.
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76. Byers, J.A. 2001. Correlated random walk equations of animal dispersal
resolved by simulation. Ecology 82:1680-1690.
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77. Byers, J.A. 2002. Internet programs for drawing moth pheromone analogs
and searching literature database. Journal of Chemical Ecology 28:
807-817. 
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78. Byers, J.A. 2004a. Equations for nickel-chromium wire heaters of column transfer
lines in gas chromatographic-electroantennographic detection (GC-EAD).
Journal of Neuroscience Methods 135:89-93.
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79. Byers, J.A., Zhang, Q.H., and Birgersson, G. 2004. Avoidance of nonhost plants
by a bark beetle, Pityogenes bidentatus, in a forest of odors.
Naturwissenschaften 91:215-219.
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80. Byers, J.A. 2004b. Chemical ecology of bark beetles in a complex olfactory landscape.
pp. 89-134. In, Bark and Wood Boring Insects in Living Trees in Europe, a Synthesis,
F. Lieutier, K.R. Day, A. Battisti, J.C. Grégoire, H. Evans (Eds). Kluwer Academic
Publishers, Dordrecht, The Netherlands.
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81. Blackmer, J.L., Rodriquez-Saona, C., Byers, J. A., Shope, K.L., and Smith, J.P. 2004.
Behavioral response of Lygus hesperus to conspecifics and headspace volatiles
of alfalfa in a Y-tube olfactometer. Journal of Chemical Ecology 30:1547-1564.
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82. Byers, J.A. 2005a. A cost of alarm pheromone production in cotton aphids, Aphis gossypii.
Naturwissenschaften 92:69-72.
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83. Byers, J.A. 2005b. Chemical constraints on the evolution of olfactory communication
channels of moths. Journal of Theoretical Biology 235:199-206.
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84. Byers, J.A. and Castle, S.J. 2005. Areawide models comparing synchronous versus
asynchronous treatments for control of dispersing insect pests. Journal of
Economic Entomology 98:1763-1773.
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85. Byers, J.A. 2006a. Pheromone component patterns of moth evolution revealed by computer
analysis of the Pherolist. Journal of Animal Ecology 75:399-407.
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86. Thoss, V., and Byers, J.A. 2006. Monoterpene chemodiversity of ponderosa pine
in relation to herbivory and bark beetle colonization. Chemoecology 16:51-58.
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87. Byers, J.A. 2006b. Analysis of insect and plant colors in digital images using
Java software on the Internet. Annals of the Entomological Society of America
99:865-874.
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88. El-Sayed, A.M., Suckling, D.M., Wearing, C.H., and Byers, J.A. 2006. Potential of
mass trapping for long-term pest management and eradication of invasive species.
Journal of Economic Entomology 99:1550-1564.
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89. Byers, J.A. 2006c. Production and predator-induced release of volatile chemicals by
the plant bug Lygus hesperus. Journal of Chemical Ecology 32:2205-2218.
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90. Byers, J.A. 2007. Simulation of mating disruption and mass trapping with competitive
attraction and camouflage. Environmental Entomology 36:1328-1338.
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91. Natwick, E.T, Byers, J.A., Chu, C.C., Lopez, M., and Henneberry, T.J. 2007. Early
detection and mass trapping of Frankliniella occidentalis and Thrips tabaci
in vegetable crops. Southwestern Entomologist 32:229-238.
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92. Blackmer, J.L., Byers, J.A., and Rodriguez-Saona, C. 2008. Evaluation of color traps
for monitoring Lygus spp.: Design, placement, height, time of day, and non-target
effects. Crop Protection 27:171-181.
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93. El-Sayed, A.M, Byers, J.A., Manning, L.M., Jürgens, A., Mitchell, V.J., and Suckling,
D.M. 2008. Floral scent of Canada thistle, Cirsium arvense (L.) Scop and its
potential as a generic insect attractant. Journal of Economic Entomology in press.
Popular Science and Education:
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My web site previewed as "Bug juice" in Science 288:767. on 5 May 2000. |
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Theses:
Byers, J.A. 1973. Plant growth hormones in pinyon insect galls. Master
of Science, Department of Entomology, Colorado State University,
Fort Collins, Colorado, U.S.A.
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Byers, J.A. 1978. Mutual inhibition of the attractive response to
pheromones of Dendroctonus brevicomis and Ips paraconfusus, and the
physiology of pheromone production in Ips paraconfusus. Doctor of
Philosophy, Department of Entomological Sciences, University of
California, Berkeley, California, U.S.A.
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Miscellaneous Letters in Entomological Publications:
Byers, J.A. 1983. (Letter proposing new journal called `Entomology'). ESA
Newsletter (Entomological Society of America) 6(10):1.
Byers, J.A. 1984. (Letter proposing name change of `Ann. Entomol. Soc.
Am.' to `Entomology', also proposes free page charges to increase
competition and quality in ESA journals). ESA Newsletter 7(8):6.
Byers, J.A. 1985. (Letter discussing International membership, a recent
questionnaire, `Entomology' journal, and the need for having
national meetings in different regions). ESA Newsletter 8(10):3.
Byers, J.A. 1987. What's in a name? Bulletin of the Entomological Society
of America 33:208.